Predicted to enable proteasome-activating activity. Predicted to be involved in proteasome-mediated ubiquitin-dependent protein catabolic process. Predicted to be part of proteasome regulatory particle, base subcomplex. Is expressed in central brain primordium; embryonic brain; fat body/gonad primordium; gonad; and organism. Orthologous to human PSMC4 (proteasome 26S subunit, ATPase 4).